6W. Panthera (Leo) spelaea, the cave lion, is the one in the middle; males did not have manes. (If you want to make your head hurt, dive into the discussion boards about relative sizes of various felines, past and present.) -- In "Clan...." we meet the cave lion early on and his presence is felt throughout, as he comes to symbolize Ayla in the way the cave bear does for the clan.
Panthera leo spelaea (cave lion) - recent genetic research shows that it was most closely related to the modern lion and that it formed a single population with the Beringian cave lion. It ranged from Europe to Alaska over the Bering land bridge until the late Pleistocene.
8W. Photo: "Panthera-Leo-spelaea" -- [Creb said,] “ 'Spirit of Cave Lion, the girl, Ayla, is delivered into your protection.' ...The idea of taking the strange child into the clan had been difficult enough, but this totem of hers was too much. It was irregular, unconventional; Brun didn’t like anomalies in his well-ordered clan... There would be no further deviations. If the girl was to be a member of his clan, she would conform, Cave Lion or no Cave Lion." p.89-90